Koko, the gorilla, who learned sign language is no more

46-year-old Koko, the gorilla who was an expert in sign language, has died. The Gorilla Foundation in California in the United States says the western lowland gorilla died in her sleep at the foundation's sanctuary.

Koko was born at the San Francisco Zoo. She was taught gorilla sign language by an academic from the Stanford University as part of a project in 1974.

The Gorilla Foundation said Koko's capacity for language and understanding opened the minds and hearts of thousands. Koko appeared in many documentaries and twice in the National Geographic magazine.

The foundation plans to honour Koko's legacy with a sign language application featuring Koko for the benefit of gorillas and children, as well as other projects.

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